2.4.1.212 hyaluronan synthase fibroblast-like synoviocyte FLS 693068 2.7.10.2 non-specific protein-tyrosine kinase fibroblast-like synoviocyte FLS 693455 3.4.24.B4 matrix metalloproteinase-13 fibroblast-like synoviocyte from rheumatoid arthritis patients 711041